Born and raised in Connecticut, I earned BS degrees in biology and economics and a PhD in biology from Boston College in Chestnut Hill, MA. I next moved to Houston, Texas for a postdoctoral fellowship and research assistant professor appointment in the Department of Physiology and Cell Biology at the University of Texas Health Science Center-Houston. In Houston I worked on projects related to cystic fibrosis. Our family relocated to Johnson City, TN when I joined the faculty of the Quillen College of Medicine in 1995. My basic research interest is the cell biology of ion transporting epithelia and my current focus is the evolution of calcium transport mechanisms in the extraembryonic membranes of oviparous and viviparous reptiles. Since joining QCOM I have taught the gastrointestinal section of the Medical Physiology course for first year students. Since 2012 I have served as director of that course and have expanded my teaching duties to include basic cellular and reproductive physiology. I also serve as a facilitator in the Case-Based Learning section of Doctoring 1. My outside interests include hiking, camping, and trail ultra-marathons.